九龙山紫菀
——浙江紫菀属(菊科)一新种
2017-02-24叶喜阳陈征海潘成椿马丹丹李根有
叶喜阳, 陈征海, 潘成椿, 马丹丹, 李根有
(1. 浙江农林大学林业与生物技术学院,浙江 临安 311300; 2. 浙江省森林资源监测中心,浙江 杭州 310020; 3. 浙江九龙山国家级自然保护区,浙江 遂昌 323301; 4. 浙江农林大学暨阳学院,浙江 诸暨 311800)
九龙山紫菀
——浙江紫菀属(菊科)一新种
叶喜阳1, 陈征海2, 潘成椿3, 马丹丹4, 李根有4
(1. 浙江农林大学林业与生物技术学院,浙江 临安 311300; 2. 浙江省森林资源监测中心,浙江 杭州 310020; 3. 浙江九龙山国家级自然保护区,浙江 遂昌 323301; 4. 浙江农林大学暨阳学院,浙江 诸暨 311800)
描述了浙江九龙山菊科Asteraceae紫菀属Aster一新种:九龙山紫菀Asterjiulongshanensis,并附有墨线图.新种接近于长叶紫菀A.dolichophyllus,但其茎中部以上多分枝,基部及下部的叶花期枯萎,全部叶片被毛,厚纸质,羽状脉,复合花序由7~10个伞房状排列的狭圆锥状支花序组成,头状花序直径约1.5 cm,总苞狭倒锥形或管状,总苞片3或4层,先端不外展,舌状花5~10个,舌片绿白色或白色;冠毛等长,白色或污白色而明显有别.
九龙山紫菀;紫菀属;菊科;新种;浙江
近年来,我们在野生植物资源调查研究和《浙江植物志》(第二版)的编写过程中,在浙江西南部山区发现了一种紫菀属植物,经查阅文献资料[1-8]和栽培观察研究,确认为一新种,现予报道.
九龙山紫菀 新种(图1、图2)
Aster jiulongshanensis Z. H. Chen, X. Y.Ye et C .C. Pan, sp. nov.(Fig.1,2)
The new species is closed toA.dolichophyllus, but differs in having the ramose multiple, from middle part of stem; leaves pubescent, thickly papery, pinnately veined, basal and lowermost ones wither in blooming period; synflorescens with 7~10 narrow panicles, arranged in corymbiform sub-synflorescens; capitulum approximately 1.5 cm in diam.; involucres narrowly obconic or tubiform; phyllaris 3 or 4 seriate, apex not expanding; ligulate flowers 5 to 10; ray florets greenish white or white; pappus equal in length, white or beige.
Type: China, Zhejiang (浙江), Suichang (遂昌), Mount. Jiulong (九龙山), Huangtanyu (黄坛淤),on rocks in valley, alt. 530 m, 30 Oct. 2011,G.Y.LietZ.H.Chen(李根有, 陈征海)JLS20111001 (holotype, ZJFC).
图1 九龙山紫菀(Ⅰ)Fig. 1 Aster jiulongshanensis(Ⅰ)
1 总苞与总苞片; 2 总苞片; 3 舌状花; 4 管状花; 5 雄蕊与雌蕊(管状花); 6 瘦果及冠毛.图2 九龙山紫菀(Ⅱ)Fig. 2 Aster jiulongshanensis(Ⅱ)
多年生草本.根状茎粗壮,匍匐枝长8~12 cm,稍木质.茎直立或斜升,高50~130 cm,中部以上多分枝,被卷曲的糙毛或下部近光滑.基生叶密集成莲座状,在花期枯萎,叶片长卵形至长卵状披针形,长4~13 cm,宽2~9 cm,先端渐尖或急尖,基部楔形下延成长柄;中部叶疏生,叶片卵状披针形,长7~12 cm,宽2~4 cm,基部渐狭下延至叶柄;上部叶片渐小,倒披针形,长2~4 cm,宽0.8~2 cm;全部叶片厚纸质,正面被糙毛,下部叶尤密,背面无毛或沿脉具稀疏毛,边缘密被开展的糙毛,有疏锯齿,齿端具外展的小尖头,羽状脉,侧脉3~5对,近叶缘处网结;叶柄具翅,基生叶的叶柄长4~8(~16)cm,基部扩大半抱茎,茎中、上部的叶柄渐短.复合花序由7~10个伞房状排列的支花序组成,每个支花序呈狭圆锥状或总状,长15~38 cm,由5~10(~32)个总状花序组成,每个总状花序具1~5个头状花序;头状花序直径约1.5 cm;花序梗密被毛,苞叶长圆状披针形,全缘;总苞狭倒锥形或管状,基部直径0.5~0.6 cm,长1~1.2 cm;总苞片3或4层,条形至条状披针形,草质,边缘膜质,背部绿色,先端渐尖,外层较短,长约2 mm,具缘毛,内层较长,5~6 mm,中脉草质,沿脉疏被毛,最内层仅先端具少量缘毛.舌状花5~10个,1层,舌片绿白色或白色,条形,长约4 mm,宽约1 mm;管状花多数,长5~6 mm,管部长2~3 mm,被柔毛,裂片卷曲;花柱附片长披针形.瘦果条状披针形,稍扁,长约3.5 mm,宽约0.75 mm,熟时由紫红色转褐色,有2条边肋,两面各有1条中肋,被上向白色短糙毛;冠毛等长,长5~6 mm,白色或污白色,不分枝,具向上短糙毛.花果期9~11月.
Paratypes: Zhejiang (浙江), Suichang (遂昌), Mount. Jiulong (九龙山), Huangtanyu (黄坛淤), roadside in valley, alt. 450 m, 26 Oct. 2012,Z.H.ChenetX.Y.Ye(陈征海, 叶喜阳)JLS2012001 (ZJFC); the same locality, alt. 530 m, 26 Oct. 2012,Z.H.ChenetX.Y.Ye(陈征海, 叶喜阳)JLS2012004 (ZJFC). Jiangshan (江山), Nianbadu (廿八都), Zhoucun (周村), Lidongkeng (里东坑), wet rocky slope of a valley at forest margin, alt. 600 m, 28 Sep. 2015,Z.H.Chen,W.Y.XieetF.Y.Zhang(陈征海, 谢文远, 张芬耀)JS150903 (ZJFC). Jingning (景宁), Yingchuan (英川), Mudaikou (木岱口), rocky slope at forest margin, alt. 680 m, 2 Nov. 2016,Z.H.ChenetG.Y.Li(陈征海, 李根有)JN20161102 (ZJFC). Longquan (龙泉),Mount. Maoshan (峁山), rocky slope at forest margin, alt. 520 m, 11 Sept. 2013,Z.H.ChenetL.M.Ji(陈征海, 季利民)s.n. (ZJFC). Qingyuan (庆元), Qiangangkeng (千岗坑), wet rocky slope under forest, alt. 820 m, 1 Aug. 2014,Z.H.ChenetG.Y.Li(陈征海, 李根有)s.n. (ZJFC).
致谢 浙江农林大学夏国华副教授帮助对花部进行解剖并扫描;上海辰山植物园葛斌杰帮助对瘦果和冠毛进行电镜扫描;张逸帮助绘制线描图;杭州师范大学金孝锋教授对初稿进行审阅并提出修改意见,在此一并致以诚挚的谢意!
[1] 林镕.紫菀属(菊科)[M]//林镕,陈艺林,石铸.中国植物志:第七十四卷.北京:科学出版社,1985:133-254.
[2] CHEN Y L, BROUILLET L, SEMPLE J C.AsterLinn[M]// WU Z Y, RAVEN P H. Flora of China, Vol.20-21. Beijing: Science Press & St. Louis: Missouri Botanical Garden Press, St. Louis,2011:574-632.
[3] 郑朝宗.菊科[M]//郑朝宗.浙江植物志:第六卷.杭州:浙江科学技术出版社,1993:212-368.
[4] 陈丽云,林来官.菊科[M]//福建省科学技术委员会,《福建植物志》编写组.福建植物志:第五卷.福州:福建科学技术出版社,1993:278-447.
[5] 《安徽植物志》协作组.菊科[M]//安徽植物志:第四卷.合肥:安徽科学技术出版社,1991:476-641.
[6] 郑朝宗.浙江种子植物检索鉴定手册[M].杭州:浙江科学技术出版社,2005:385-386.
[7] SOEJIMA A, PENG C Ⅰ.AsterLinn[M]//HUANG T C. Flora of Taiwan, Vol.4. 2nd. Taipei: Editorial Committee of the Flora of Taiwan,1998:848-868.
[8] OHWI J. Flora of Japan[M]. Washington: Smthsonian Institution,1965: 872-876.
AsterJiulongshanensis(Asteraceae), a New Species from Zhejiang
YE Xiyang1, CHEN Zhenghai2, PAN Chengchun3, MA Dandan4, LI Genyou4
(1. School of Forestry and Biotechnology, Zhejiang A& F University, Lin’an 311300, China; 2. Zhejiang Monitoring Center of Forest Resources, Hangzhou 310020, China; 3. Administration of Jiulongshan National Natural Reserve, Suichang 323300, China;4. Jiyang College, Zhejiang A & F University,Zhuji 311800, China)
Asterjiulongshanensis, a new species of Asteraceae from Mount. Jiulong of Suichang, is described with illustrations. This new species is similar toA.dolichophyllus, but differs in having multiple branches from middle part of stem, leaves pubescent, thickly papery, pinnately veined, basal and lowermost ones wither in blooming period, synflorescens with 7~10 narrow panicles, arranged in corymbiform sub-synflorescens, capitulum approximately 1.5 cm in diam., involucres narrowly obconic or tubiform, phyllaris 3 or 4 seriate, apex not expanding, ligulate flowers 5 to 10, ray florets greenish white or white, pappus equal in length, white or beige.
Asterjiulongshanensis;Aster; Asteraceae; new species; Zhejiang
2016-09-10
国家林业局第二次全国重点保护野生植物资源调查项目(林护发[2012]87号); 浙江省第二次重点保护野生植物资源调查项目(335006-2013-0001); 浙江省野生植物资源调查、建档、编纂及《浙江植物志》(第二版)编著项目(335010-2015-0005).
李根有(1956—), 男,教授,主要从事植物分类与资源植物开发利用研究. E-mail: ligy1956@163.com
10.3969/j.issn.1674-232X.2017.01.004
Q949
A
1674-232X(2017)01-0013-03